About the Authors

Hiroki Moribe

himoribe@med.kurume-u.ac.jp

Affiliations Department of Biology, Kurume University School of Medicine, Kurume, Fukuoka, Japan, Department of Cell Biology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Ryouji Konakawa

Affiliation Department of Cell Biology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Daisuke Koga

Affiliation Division of Microscopic Anatomy and Bio-Imaging, Niigata University Graduate School of Medical and Dental Sciences, Chuo-ku, Niigata, Japan

Tatsuo Ushiki

Affiliation Division of Microscopic Anatomy and Bio-Imaging, Niigata University Graduate School of Medical and Dental Sciences, Chuo-ku, Niigata, Japan

Kuniaki Nakamura

Affiliation Department of Cell Biology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Eisuke Mekada

Affiliation Department of Cell Biology, Research Institute for Microbial Diseases, Osaka University, Suita, Osaka, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HM EM. Performed the experiments: HM RK DK KN. Analyzed the data: HM. Contributed reagents/materials/analysis tools: HM RK DK TU. Wrote the paper: HM.