About the Authors

Keiko Myouzen, Yuta Kochi, Yukinori Okada, Akari Suzuki, Kazuhiko Yamamoto
Laboratory for Autoimmune Diseases, Center for Genomic Medicine (CGM), RIKEN, Yokohama, Japan

Yuta Kochi, Yukinori Okada, Kazuhiko Yamamoto
Department of Allergy and Rheumatology, Graduate School of Medicine, the University of Tokyo, Tokyo, Japan

Yukinori Okada, Atsushi Takahashi
Laboratory for Statistical Analysis, CGM, RIKEN, Yokohama, Japan

Chikashi Terao, Fumihiko Matsuda
Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan

Chikashi Terao, Koichiro Ohmura, Tsuneyo Mimori
Department of Rheumatology and Clinical Immunology, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Katsunori Ikari, Atsuo Taniguchi, Shigeki Momohara, Hisashi Yamanaka
Institute of Rheumatology, Tokyo Women's Medical University, Tokyo, Japan

Tatsuhiko Tsunoda
Laboratory for Medical Informatics, CGM, RIKEN, Yokohama, Japan

Michiaki Kubo
Laboratory for Genotyping Development, CGM, RIKEN, Yokohama, Japan

Fumihiko Matsuda
CREST Program, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan

Fumihiko Matsuda
Institut National de la Sant? et de la Recherche M?dicale (INSERM), Unit? U852, Kyoto University Graduate School of Medicine, Kyoto, Japan

Naoyuki Kamatani
Laboratory for International Alliance, CGM, RIKEN, Yokohama, Japan

Ryo Yamada
Unit of Statistical Genetics, Center for Genomic Medicine, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Yusuke Nakamura
Laboratory of Molecular Medicine, Human Genome Center, Institute of Medical Science, University of Tokyo, Tokyo, Japan

Corresponding Author

ykochi@src.riken.jp

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: K Myouzen, Y Kochi, Y Okada, C Terao, K Ikari, K Ohmura, R Yamada, K Yamamoto. Performed the experiments: K Myouzen, Y Kochi, C Terao, A Suzuki, K Ikari, K Ohmura. Analyzed the data: K Myouzen, Y Kochi, Y Okada, C Terao, T Tsunoda, A Takahashi, R Yamada. Contributed reagents/materials/analysis tools: M Kubo, A Taniguchi, F Matsuda, K Ohmura, S Momohara, T Mimori, H Yamanaka, N Kamatani, Y Nakamura. Wrote the paper: K Myouzen, Y Kochi, Y Okada, C Terao, K Yamamoto.