About the Authors

Fan Liu, Fedde van der Lijn, Andreas Wollstein, Oscar Lao, Manfred Kayser
Department of Forensic Molecular Biology, Erasmus MC University Medical Center Rotterdam, Rotterdam, The Netherlands

Fedde van der Lijn, Marleen de Bruijne, Wiro J. Niessen
Department of Medical Informatics, Erasmus MC University Medical Center Rotterdam, Rotterdam, The Netherlands

Fedde van der Lijn, Marleen de Bruijne, M. Arfan Ikram, Aad van der Lugt, Wiro J. Niessen
Department of Radiology, Erasmus MC University Medical Center Rotterdam, Rotterdam, The Netherlands

Claudia Schurmann, Georg Homuth
Interfaculty Institute for Genetics and Functional Genomics, Ernst-Moritz-Arndt University Greifswald, Greifswald, Germany

Gu Zhu, Sarah E. Medland, Grant W. Montgomery, Margaret J. Wright, Nicholas G. Martin
Queensland Institute of Medical Research, Brisbane, Australia

M. Mallar Chakravarty, Tomáš Paus
Rotman Research Institute, University of Toronto, Toronto, Canada

M. Mallar Chakravarty
Centre for Addiction and Mental Health, University of Toronto, Toronto, Canada

Pirro G. Hysi, Liisa Bevan, Timothy D. Spector
Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

M. Arfan Ikram, Fernando Rivadeneira, André G. Uitterlinden, Albert Hofman
Department of Epidemiology, Erasmus MC University Medical Center Rotterdam, Rotterdam, The Netherlands

Fernando Rivadeneira, André G. Uitterlinden
Department of Internal Medicine, Erasmus MC University Medical Center Rotterdam, Rotterdam, The Netherlands

Wiro J. Niessen
Department of Imaging Science and Technology, Faculty of Applied Sciences, Delft University of Technology, Delft, The Netherlands

Greig de Zubicaray, Katie L. McMahon
Centre for Advanced Imaging, University of Queensland, Brisbane, Australia

Paul M. Thompson
Laboratory of Neuroimaging, School of Medicine, University of California Los Angeles, Los Angeles, California, United States of America

Amro Daboul, Reiner Biffar
Center of Oral Health, Department of Prosthodontics, Gerostomatology, and Dental Materials, University Medicine Greifswald, Greifswald, Germany

Ralf Puls, Katrin Hegenscheid
Department of Radiology and Neuroradiology, University Medicine Greifswald, Greifswald, Germany

Zdenka Pausova
The Hospital of Sick Children, University of Toronto, Toronto, Canada

Carol Wicking
Institute for Molecular Bioscience, University of Queensland, Brisbane, Australia

Stefan Boehringer
Department of Medical Statistics and Bioinformatics, Leiden University Medical Center, Leiden, The Netherlands

Tomáš Paus
Montréal Neurological Institute, McGill University, Montréal, Canada

Corresponding Author

m.kayser@erasmusmc.nl

Competing Interests

TS and MK have consulted for Identitas Inc. and are on the SAB but without financial or other direct benefits. All other authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FL MK. Analyzed the data: FL FvdL CS GZ MMC AW OL SB. Contributed reagents/materials/analysis tools: MK RB NGM TP TDS AH AGU MAI. Data preparation: FvdL PGH MdB AW MAI AvdL FR AGU AH WJN GH GdZ KLM PMT AD RP KH LB ZP SEM GWM MJW TDS TP NGM RB. Wrote the manuscript: FL MK. Additional contributions: CW FvdL AW NGM CS MMC OL MAI SB TDS TP RB. Approved the manuscript: FL FvdL CS GZ MMC PGH AW OL MdB MAI AvdL FR AGU AH WJN GH GdZ KLM PMT AD RP KH LB ZP SEM GWM MJW CW SB TDS TP NGM RB MK.