About the Authors

Jennifer S. Yokoyama, Carolyn A. Erdman, Steven P. Hamilton
Department of Psychiatry and Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Ernest T. Lam
Institute for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Alison L. Ruhe, Mark W. Neff
Neurogenomics Division, The Translational Genomics Research Institute, Phoenix, Arizona, United States of America

Alison L. Ruhe, Kathryn R. Robertson
Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Aubrey A. Webb
CullenWebb Animal Neurology and Ophthalmology Centre, Riverview, New Brunswick, Canada

Aubrey A. Webb
Department of Clinical Neuroscience, Faculty of Medicine, University of Calgary, Calgary, Alberta, Canada

D. Colette Williams
School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Melanie L. Chang
Department of Anthropology, University of Oregon, Eugene, Oregon, United States of America

Marjo K. Hytönen, Hannes Lohi
Department of Veterinary Biosciences and Research Programs Unit, Molecular Medicine, University of Helsinki and Folkhälsan Research Center, Helsinki, Finland

Mark W. Neff
The Van Andel Research Institute, Grand Rapids, Michigan, United States of America

Corresponding Author

mark.neff@vai.org

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JSY ETL SPH MWN. Performed the experiments: JSY ETL ALR CAE KRR MKH. Analyzed the data: JSY ETL. Contributed reagents/materials/analysis tools: JSY ETL ALR CAE KRR AAW DCW MLC MKH HL SPH MWN. Wrote the paper: JSY ETL CAE AAW MLC SPH MWN. Critical reading and discussion of manuscript: AAW DCW HL ALR.