About the Authors

Madhura Kulkarni
Department of Cell Biology and Molecular Genetics, University of Maryland, College Park, Maryland, United States of America

Diane C. Shakes, Katie Guevel
Department of Biology, College of William and Mary, Williamsburg, Virginia, United States of America

Harold E. Smith
National Institute of Diabetes and Digestive and Kidney Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Corresponding Author

smithhe2@niddk.nih.gov

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HES DCS. Performed the experiments: MK DCS KG HES. Analyzed the data: MK DCS KG HES. Contributed reagents/materials/analysis tools: MK DCS HES. Wrote the paper: MK DCS HES.