About the Authors

Artyom A. Alekseyenko

Contributed equally to this work with: Artyom A. Alekseyenko, Joshua W. K. Ho, Shouyong Peng

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Joshua W. K. Ho

Contributed equally to this work with: Artyom A. Alekseyenko, Joshua W. K. Ho, Shouyong Peng

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Center for Biomedical Informatics, Harvard Medical School, Boston, Massachusetts, United States of America

Shouyong Peng

Contributed equally to this work with: Artyom A. Alekseyenko, Joshua W. K. Ho, Shouyong Peng

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Center for Biomedical Informatics, Harvard Medical School, Boston, Massachusetts, United States of America

Marnie Gelbart

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Michael Y. Tolstorukov

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Center for Biomedical Informatics, Harvard Medical School, Boston, Massachusetts, United States of America

Annette Plachetka

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Peter V. Kharchenko

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Center for Biomedical Informatics, Harvard Medical School, Boston, Massachusetts, United States of America

Youngsook L. Jung

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Center for Biomedical Informatics, Harvard Medical School, Boston, Massachusetts, United States of America

Andrey A. Gorchakov

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Erica Larschan

Affiliation Department of Molecular Biology, Cell Biology, and Biochemistry, Brown University, Providence, Rhode Island, United States of America

Tingting Gu

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Aki Minoda

Affiliations Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America, Department of Genome Dynamics, Lawrence Berkeley National Lab, Berkeley, California, United States of America

Nicole C. Riddle

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Yuri B. Schwartz

Affiliation Department of Molecular Biology, Umeå University, Umeå, Sweden

Sarah C. R. Elgin

Affiliation Department of Biology, Washington University in St. Louis, St. Louis, Missouri, United States of America

Gary H. Karpen

Affiliation Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America

Vincenzo Pirrotta

Affiliation Department of Molecular Biology and Biochemistry, Rutgers University, Piscataway, New Jersey, United States of America

Mitzi I. Kuroda

mkuroda@genetics.med.harvard.edu (MIK); peter_park@harvard.edu (PJP)

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Peter J. Park

mkuroda@genetics.med.harvard.edu (MIK); peter_park@harvard.edu (PJP)

Affiliations Division of Genetics, Department of Medicine, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Center for Biomedical Informatics, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: GHK SCRE VP PJP MIK. Performed the experiments: AAA MG AP AAG EL TG AM NCR YBS. Analyzed the data: JWKH SP MYT PVK YLJ PJP. Wrote the paper: JWKH AAA PJP MIK.