About the Authors

Barbara E. Stranger

Contributed equally to this work with: Barbara E. Stranger, Stephen B. Montgomery, Antigone S. Dimas

Affiliations Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom, Department of Medicine, Division of Genetics, Brigham and Women's Hospital, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Harvard Medical School, Boston, Massachusetts, United States of America

Stephen B. Montgomery

Contributed equally to this work with: Barbara E. Stranger, Stephen B. Montgomery, Antigone S. Dimas

Affiliations Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom, Department of Genetic Medicine and Development, University of Geneva Medical School, Geneva, Switzerland, Institute of Genetics and Genomics in Geneva (iGE3), Geneva, Switzerland

Antigone S. Dimas

Contributed equally to this work with: Barbara E. Stranger, Stephen B. Montgomery, Antigone S. Dimas

Affiliations Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom, Department of Genetic Medicine and Development, University of Geneva Medical School, Geneva, Switzerland, Institute of Genetics and Genomics in Geneva (iGE3), Geneva, Switzerland, Wellcome Trust Centre for Human Genetics, Oxford, United Kingdom, Max Planck Institute Intelligent Systems and Max Planck Institute for Developmental Biology, Tübingen, Germany

Leopold Parts

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Oliver Stegle

Affiliation Max Planck Institute Intelligent Systems and Max Planck Institute for Developmental Biology, Tübingen, Germany

Catherine E. Ingle

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Magda Sekowska

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

George Davey Smith

Affiliation MRC CAiTE centre, School of Social and Community Medicine, University of Bristol, Bristol, United Kingdom

David Evans

Affiliation MRC CAiTE centre, School of Social and Community Medicine, University of Bristol, Bristol, United Kingdom

Maria Gutierrez-Arcelus

Affiliations Department of Genetic Medicine and Development, University of Geneva Medical School, Geneva, Switzerland, Institute of Genetics and Genomics in Geneva (iGE3), Geneva, Switzerland

Alkes Price

Affiliations Program in Medical and Population Genetics, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Department of Epidemiology and Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America

Towfique Raj

Affiliations Department of Medicine, Division of Genetics, Brigham and Women's Hospital, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of Harvard and Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America, Department of Neurology, Brigham and Women's Hospital, Boston, Massachusetts, United States of America

James Nisbett

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Alexandra C. Nica

Affiliations Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom, Department of Genetic Medicine and Development, University of Geneva Medical School, Geneva, Switzerland, Institute of Genetics and Genomics in Geneva (iGE3), Geneva, Switzerland

Claude Beazley

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Richard Durbin

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Panos Deloukas

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom

Emmanouil T. Dermitzakis

emmanouil.dermitzakis@unige.ch

Affiliations Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, United Kingdom, Department of Genetic Medicine and Development, University of Geneva Medical School, Geneva, Switzerland, Institute of Genetics and Genomics in Geneva (iGE3), Geneva, Switzerland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BES SBM ASD PD ETD. Performed the experiments: CEI MS JN. Analyzed the data: BES SBM ASD LP OS ACN CB TR MG-A. Contributed reagents/materials/analysis tools: GDS DE RD AP. Wrote the paper: BES SBM ASD ETD.