About the Authors

Frederick J. Tan

Contributed equally to this work with: Frederick J. Tan, Margaret L. Hoang

Affiliations Howard Hughes Medical Institute and Department of Embryology, Carnegie Institution, Baltimore, Maryland, United States of America, Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America

Margaret L. Hoang

Contributed equally to this work with: Frederick J. Tan, Margaret L. Hoang

Current address: Ludwig Center for Cancer Genetics and Therapeutics, Johns Hopkins Medical Institute, Baltimore, Maryland, United States of America

Affiliations Howard Hughes Medical Institute and Department of Embryology, Carnegie Institution, Baltimore, Maryland, United States of America, Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States of America

Douglas Koshland

koshland@berkeley.edu

Affiliations Howard Hughes Medical Institute and Department of Embryology, Carnegie Institution, Baltimore, Maryland, United States of America, Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FJT MLH DK. Performed the experiments: FJT MLH. Analyzed the data: FJT MLH DK. Contributed reagents/materials/analysis tools: FJT MLH. Wrote the paper: FJT MLH DK.