About the Authors

Shanshan Zhou

Affiliations Department of Biology, North Carolina State University, Raleigh, North Carolina, United States of America, W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America

Terry G. Campbell

Affiliations W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America, Department of Genetics, North Carolina State University, Raleigh, North Carolina, United States of America

Eric A. Stone

Affiliations W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America, Department of Genetics, North Carolina State University, Raleigh, North Carolina, United States of America

Trudy F. C. Mackay

Affiliations W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America, Department of Genetics, North Carolina State University, Raleigh, North Carolina, United States of America

Robert R. H. Anholt

anholt@ncsu.edu

Affiliations Department of Biology, North Carolina State University, Raleigh, North Carolina, United States of America, W. M. Keck Center for Behavioral Biology, North Carolina State University, Raleigh, North Carolina, United States of America, Department of Genetics, North Carolina State University, Raleigh, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SZ TFCM RRHA. Performed the experiments: SZ TGC. Analyzed the data: SZ EAS. Wrote the paper: SZ TFCM RRHA.