About the Authors

Xiaolin Tu

Address Current address: Department of Anatomy and Cell Biology, Indiana University School of Medicine, Indianapolis, Indiana, United States of America

Affiliation Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America

Jianquan Chen

Affiliation Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America

Joohyun Lim

Affiliation Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America

Courtney M. Karner

Affiliation Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America

Seung-Yon Lee

Affiliation Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America

Julia Heisig

Affiliation Developmental Biochemistry, Biocentre, University of Wuerzburg, Wuerzburg, Germany

Cornelia Wiese

Affiliation Developmental Biochemistry, Biocentre, University of Wuerzburg, Wuerzburg, Germany

Kameswaran Surendran

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Raphael Kopan

Affiliation Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Manfred Gessler

Affiliation Developmental Biochemistry, Biocentre, University of Wuerzburg, Wuerzburg, Germany

Fanxin Long

flong@wustl.edu

Affiliations Department of Medicine, Washington University School of Medicine, St. Louis, Missouri, United States of America, Department of Developmental Biology, Washington University School of Medicine, St. Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FL. Performed the experiments: XT JC JL CMK S-YL JH CW. Analyzed the data: XT FL. Contributed reagents/materials/analysis tools: KS RK MG. Wrote the paper: XT FL.