About the Authors

Daisuke Sakai

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America, Graduate School of Biological Science, Nara Institute of Science and Technology, Ikoma, Japan

Jill Dixon

Affiliation Faculty of Medical and Human Sciences, Manchester Academic Health Sciences Centre, University of Manchester, Manchester, United Kingdom

Michael J. Dixon

Affiliations Faculty of Medical and Human Sciences, Manchester Academic Health Sciences Centre, University of Manchester, Manchester, United Kingdom, Faculty of Life Sciences, University of Manchester, Manchester, United Kingdom

Paul A. Trainor

pat@stowers.org

Affiliations Stowers Institute for Medical Research, Kansas City, Missouri, United States of America, Department of Anatomy and Cell Biology, University of Kansas Medical Center, Kansas City, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DS PAT. Performed the experiments: DS. Analyzed the data: DS PAT. Contributed reagents/materials/analysis tools: JD MJD. Wrote the paper: DS JD MJD PAT.