About the Authors

Song Liu

Contributed equally to this work with: Song Liu, Tomoyo Sawada

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Tomoyo Sawada

Contributed equally to this work with: Song Liu, Tomoyo Sawada

Affiliations Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan, CREST (Core Research for Evolutionary Science and Technology), Japan Science and Technology Agency, Saitama, Japan

Seongsoo Lee

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Wendou Yu

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

George Silverio

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Philomena Alapatt

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Ivan Millan

Affiliation Neuroscience Program, Stanford University School of Medicine, Stanford, California, United States of America

Alice Shen

Affiliation College of Arts and Sciences, Washington University, St. Louis, Missouri, United States of America

William Saxton

Affiliation Department of Molecular, Cell, and Developmental Biology, University of California Santa Cruz, Santa Cruz, California, United States of America

Tomoko Kanao

Affiliation Research Institute for Diseases of Old Age, Juntendo University Graduate School of Medicine, Tokyo, Japan

Ryosuke Takahashi

Affiliations Department of Neurology, Graduate School of Medicine, Kyoto University, Kyoto, Japan, CREST (Core Research for Evolutionary Science and Technology), Japan Science and Technology Agency, Saitama, Japan

Nobutaka Hattori

Affiliations CREST (Core Research for Evolutionary Science and Technology), Japan Science and Technology Agency, Saitama, Japan, Department of Neurology, Juntendo University Graduate School of Medicine, Tokyo, Japan

Yuzuru Imai

yzimai@juntendo.ac.jp (YI); bingwei@stanford.edu (BL)

Affiliation Department of Neuroscience for Neurodegenerative Disorders, Juntendo University Graduate School of Medicine, Tokyo, Japan

Bingwei Lu

yzimai@juntendo.ac.jp (YI); bingwei@stanford.edu (BL)

Affiliation Department of Pathology, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SL BL YI TS RT. Performed the experiments: SL TS SL WY GS PA IM AS TK YI. Analyzed the data: SL TS WS RT YI BL. Contributed reagents/materials/analysis tools: WS NH. Wrote the paper: BL SL YI.