About the Authors

Jirong Long

Contributed equally to this work with: Jirong Long, Qiuyin Cai

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt-Ingram Cancer Center, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Qiuyin Cai

Contributed equally to this work with: Jirong Long, Qiuyin Cai

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt-Ingram Cancer Center, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Hyuna Sung

Affiliation Department of Biomedical Sciences, Seoul National University College of Medicine, Seoul, Korea

Jiajun Shi

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt-Ingram Cancer Center, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Ben Zhang

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt-Ingram Cancer Center, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Ji-Yeob Choi

Affiliation Department of Biomedical Sciences, Seoul National University College of Medicine, Seoul, Korea

Wanqing Wen

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt-Ingram Cancer Center, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Ryan J. Delahanty

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt-Ingram Cancer Center, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Wei Lu

Affiliation Shanghai Center for Disease Control and Prevention, Shanghai, China

Yu-Tang Gao

Affiliation Department of Epidemiology, Shanghai Cancer Institute, Shanghai, China

Hongbing Shen

Affiliation Department of Epidemiology and Biostatistics, Nanjing Medical University, Nanjing, China

Sue K. Park

Affiliations Department of Biomedical Sciences, Seoul National University College of Medicine, Seoul, Korea, Department of Preventive Medicine, Seoul National University College of Medicine, Seoul, Korea, Cancer Research Institute, Seoul National University College of Medicine, Seoul, Korea

Kexin Chen

Affiliation Department of Epidemiology and Biostatistics, Tianjin Medical University Cancer Institute and Hospital, Tianjin, China

Chen-Yang Shen

Affiliations Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan, College of Public Health, China Medical University, Taichung, Taiwan

Zefang Ren

Affiliation School of Public Health, Sun Yat-sen University, Guangzhou, China

Christopher A. Haiman

Affiliation Department of Preventive Medicine, Keck School of Medicine, University of Southern California and Norris Comprehensive Cancer Center, Los Angeles, California, United States of America

Keitaro Matsuo

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center Research Institute, Nagoya, Japan

Mi Kyung Kim

Affiliations Cancer Epidemiology Division, Breast Cancer Center, National Cancer Center, Goyang, Korea, Department of Surgery, Asan Medical Center, University of Ulsan College of Medicine, Ulsan, Korea

Ui Soon Khoo

Affiliation Department of Pathology, Li Ka Shing Faculty of Medicine, University of Hong Kong, Hong Kong, China

Motoki Iwasaki

Affiliation Epidemiology and Prevention Division, Research Center for Cancer Prevention and Screening, National Cancer Center, Tokyo, Japan

Ying Zheng

Affiliation Shanghai Center for Disease Control and Prevention, Shanghai, China

Yong-Bing Xiang

Affiliation Department of Epidemiology, Shanghai Cancer Institute, Shanghai, China

Kai Gu

Affiliation Shanghai Center for Disease Control and Prevention, Shanghai, China

Nathaniel Rothman

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, Department of Health and Human Services, Bethesda, Maryland, United States of America

Wenjing Wang

Affiliation Shanghai Center for Disease Control and Prevention, Shanghai, China

Zhibin Hu

Affiliation Department of Epidemiology and Biostatistics, Nanjing Medical University, Nanjing, China

Yao Liu

Affiliation Department of Epidemiology and Biostatistics, Nanjing Medical University, Nanjing, China

Keun-Young Yoo

Affiliation Department of Preventive Medicine, Seoul National University College of Medicine, Seoul, Korea

Dong-Young Noh

Affiliations Cancer Research Institute, Seoul National University College of Medicine, Seoul, Korea, Department of Surgery, Seoul National University College of Medicine, Seoul, Korea

Bok-Ghee Han

Affiliation Center for Genome Science, Korea National Institute of Health, Seoul, Korea

Min Hyuk Lee

Affiliation Department of Surgery, Soonchunhyang University Hospital, Bucheon, Korea

Hong Zheng

Affiliation Department of Epidemiology and Biostatistics, Tianjin Medical University Cancer Institute and Hospital, Tianjin, China

Lina Zhang

Affiliation Department of Epidemiology and Biostatistics, Tianjin Medical University Cancer Institute and Hospital, Tianjin, China

Pei-Ei Wu

Affiliation Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan

Ya-Lan Shieh

Affiliation Institute of Biomedical Sciences, Academia Sinica, Taipei, Taiwan

Sum Yin Chan

Affiliation Department of Clinical Oncology, Queen Mary Hospital, Hong Kong, China

Shenming Wang

Affiliation First Affiliated Hospital, Sun Yat-sen University, Guangzhou, China

Xiaoming Xie

Affiliation Cancer Center, Sun Yat-sen University, Guangzhou, China

Sung-Won Kim

Affiliations Cancer Research Institute, Seoul National University College of Medicine, Seoul, Korea, Department of Surgery, Seoul National University College of Medicine, Seoul, Korea

Brian E. Henderson

Affiliation Department of Preventive Medicine, Keck School of Medicine, University of Southern California and Norris Comprehensive Cancer Center, Los Angeles, California, United States of America

Loic Le Marchand

Affiliation Epidemiology Program, Cancer Research Center, University of Hawaii, Honolulu, Hawaii, United States of America

Hidemi Ito

Affiliation Division of Epidemiology and Prevention, Aichi Cancer Center Research Institute, Nagoya, Japan

Yoshio Kasuga

Affiliation Department of Surgery, Nagano Matsushiro General Hospital, Nagano, Japan

Sei-Hyun Ahn

Affiliations Cancer Epidemiology Division, Breast Cancer Center, National Cancer Center, Goyang, Korea, Department of Surgery, Asan Medical Center, University of Ulsan College of Medicine, Ulsan, Korea

Han Sung Kang

Affiliations Cancer Epidemiology Division, Breast Cancer Center, National Cancer Center, Goyang, Korea, Department of Surgery, Asan Medical Center, University of Ulsan College of Medicine, Ulsan, Korea

Kelvin Y. K. Chan

Affiliations Department of Pathology, Li Ka Shing Faculty of Medicine, University of Hong Kong, Hong Kong, China, Department of Obstetrics and Gynecology, Li Ka Shing Faculty of Medicine, University of Hong Kong, Hong Kong, China

Hiroji Iwata

Affiliation Department of Breast Oncology, Aichi Cancer Center Central Hospital, Nagoya, Japan

Shoichiro Tsugane

Affiliation Epidemiology and Prevention Division, Research Center for Cancer Prevention and Screening, National Cancer Center, Tokyo, Japan

Chun Li

Affiliation Department of Biostatistics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Xiao-Ou Shu

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt-Ingram Cancer Center, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Dae-Hee Kang

Affiliations Department of Biomedical Sciences, Seoul National University College of Medicine, Seoul, Korea, Department of Preventive Medicine, Seoul National University College of Medicine, Seoul, Korea, Cancer Research Institute, Seoul National University College of Medicine, Seoul, Korea

Wei Zheng

wei.zheng@vanderbilt.edu

Affiliation Division of Epidemiology, Department of Medicine, Vanderbilt Epidemiology Center, Vanderbilt-Ingram Cancer Center, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the overall study: W Zheng. Performed genotyping experiments: J Shi, H Zheng. Wrote the manuscript: J Long, W Zheng, Q Cai, X-O Shu. Significantly contributed to writing the manuscript: C Li, W Wen, RJ Delahanty. Coordinated genotyping assays: Q Cai, J Long. Managed genotyping data: J Long, B Zhang. Performed statistical analyses: J Long, C Li, W Wen. Directed lab operations: Q Cai. Directed the GWAS in Korea: D-H Kang. Assisted the GWAS in Korea: H Sung, J-Y Choi. Contributed to data and biological collection of the original studies: H Shen, J-Y Choi, W Lu, Y-T Gao, H Shen, SK Park, K Chen, C-Y Shen, Z Ren, CA Haiman, K Matsuo, MK Kim, US Khoo, M Iwasaki, Y Zheng, Y-B Xiang, K Gu, N Rothman, W Wang, Z Hu, Y Liu, K-Y Yoo, D-Y Noh, B-G Han, MH Lee, H Zheng, L Zhang, P-E Wu, Y-L Shieh, SY Chan, S Wang, X Xie, S-W Kim, BE Henderson, L Le Marchand, H Ito, Y Kasuga, S-H Ahn, HS Kang, KYK Chan, H Iwata, S Tsugane, D-H Kang, X-O Shu, W Zheng.