About the Authors

Haoyan Chen

Affiliation Department of Dermatology, University of California San Francisco, San Francisco, California, United States of America

Genki Hayashi

Affiliation Department of Dermatology, University of California San Francisco, San Francisco, California, United States of America

Olivia Y. Lai

Affiliation Department of Dermatology, University of California San Francisco, San Francisco, California, United States of America

Alexander Dilthey

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

Peter J. Kuebler

Affiliation Division of Experimental Medicine, University of California San Francisco, San Francisco, California, United States of America

Tami V. Wong

Affiliation Department of Dermatology, University of California San Francisco, San Francisco, California, United States of America

Maureen P. Martin

Affiliation Cancer and Inflammation Program, Laboratory of Experimental Immunology, National Cancer Institute, Frederick, Maryland, United States of America

Marcelo A. Fernandez Vina

Affiliation Department of Pathology, Stanford University, Palo Alto, California, United States of America

Gil McVean

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

Matthias Wabl

Affiliation Department of Microbiology and Immunology, University of California San Francisco, San Francisco, California, United States of America

Kieron S. Leslie

Affiliation Department of Dermatology, University of California San Francisco, San Francisco, California, United States of America

Toby Maurer

Affiliation Department of Dermatology, University of California San Francisco, San Francisco, California, United States of America

Jeffrey N. Martin

Affiliation Department of Epidemiology and Biostatistics, University of California San Francisco, San Francisco, California, United States of America

Steven G. Deeks

Affiliation HIV/AIDS Program, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Mary Carrington

Affiliation Cancer and Inflammation Program, Laboratory of Experimental Immunology, National Cancer Institute, Frederick, Maryland, United States of America

Anne M. Bowcock

Affiliation Division of Human Genetics, Department of Genetics, Washington University School of Medicine, Saint Louis, Missouri, United States of America

Douglas F. Nixon

Affiliation Division of Experimental Medicine, University of California San Francisco, San Francisco, California, United States of America

Wilson Liao

liaowi@derm.ucsf.edu

Affiliation Department of Dermatology, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HC MW KSL TM DFN WL. Performed the experiments: MAFV AMB WL MC JNM SGD OYL GH TVW MPM. Analyzed the data: HC GH WL. Contributed reagents/materials/analysis tools: AD PJK GM. Wrote the paper: HC WL.