About the Authors

Rory Bowden

Contributed equally to this work with: Rory Bowden, Tammie S. MacFie

Affiliations Department of Statistics, University of Oxford, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Tammie S. MacFie

Contributed equally to this work with: Rory Bowden, Tammie S. MacFie

Affiliation Department of Zoology, University of Cambridge, Cambridge, United Kingdom

Simon Myers

Affiliations Department of Statistics, University of Oxford, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Broad Institute, Boston, Massachusetts, United States of America

Garrett Hellenthal

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Eric Nerrienet

Affiliation Centre Pasteur du Cameroun, Yaoundé, Cameroon

Ronald E. Bontrop

Affiliation Biomedical Primate Research Center, Rijswijk, The Netherlands

Colin Freeman

Affiliations Department of Statistics, University of Oxford, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Peter Donnelly

peter.donnelly@well.ox.ac.uk

These authors were joint senior authors on this work.

Affiliations Department of Statistics, University of Oxford, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Nicholas I. Mundy

These authors were joint senior authors on this work.

Affiliation Department of Zoology, University of Cambridge, Cambridge, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RB NIM PD TSM SM. Performed the experiments: RB TSM. Analyzed the data: RB TSM GH PD NIM SM. Contributed reagents/materials/analysis tools: NIM REB EN SM CF. Wrote the paper: RB NIM GH PD.