About the Authors

Rehan Qayyum, Lisa R. Yanek, Diane M. Becker, Lewis C. Becker
GeneSTAR Research Program, Division of General Internal Medicine, Johns Hopkins School of Medicine, Baltimore, Maryland, United States of America

Beverly M. Snively
Department of Biostatistical Sciences, Wake Forest School of Medicine, Winston-Salem, North Carolina, United States of America

Elad Ziv
Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Michael A. Nalls, Andrew B. Singleton
Laboratory of Neurogenetics, National Institute on Aging, National Institutes of Health, Bethesda, Maryland, United States of America

Yongmei Liu
Department of Epidemiology and Prevention, Division of Public Health Sciences, Wake Forest University School of Medicine, Winston-Salem, North Carolina, United States of America

Weihong Tang, Aaron R. Folsom
Division of Epidemiology and Community Health, University of Minnesota School of Public Health, Minneapolis, Minnesota, United States of America

Leslie Lange
Department of Genetics, School of Medicine, The University of North Carolina at Chapel Hill, Chapel Hill, North Carolina, United States of America

Michele K. Evans
Health Disparities Research Section, Clinical Research Branch, National Institute on Aging, National Institutes of Health, Baltimore, Maryland, United States of America

Santhi Ganesh
Division of Cardiology, University of Michigan Health System, Ann Arbor, Michigan, United States of America

Melissa A. Austin, Alexander P. Reiner
Department of Epidemiology, University of Washington, Seattle, Washington, United States of America

Melissa A. Austin
Division of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Guillaume Lettre
Montreal Heart Institute, Montreal, Canada

Alan B. Zonderman
Laboratory of Personality and Cognition, National Institute on Aging, National Institutes of Health, Baltimore, Maryland, United States of America

Tamara B. Harris
Laboratory for Epidemiology, Demography, and Biometry, National Institute on Aging, National Institutes of Health, Baltimore, Maryland, United States of America

Emile R. Mohler
Department of Medicine, University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Benjamin A. Logsdon, Charles Kooperberg
Program in Biostatistics and Biomathematics, Division of Public Health Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

James G. Wilson
Department of Medicine, University of Mississippi Medical Center, Jackson, Mississippi, United States of America

Corresponding Author

rqayyum@jhmi.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RQ BMS MAN GL MAA DMB SG ABZ ABS JGW LCB APR ERM. Performed the experiments: EZ YL MAN WT LL MKE TBH MAA DMB ABZ ABS TBH JGW LCB APR BAL ARF CK. Analyzed the data: RQ BMS LRY DMB TBH JGW APR BAL CK. Contributed reagents/materials/analysis tools: BMS EZ MAN YL WT LRY MKE SG MAA GL DMB ABZ ABS TBH JGW LCB APR ERM BAL ARF CK. Wrote the paper: RQ DMB LCB APR. Interpreted the results and revised the paper: RQ BMS EZ MAN YL WT LRY LL SG MAA GL DMB JGW LCB ARF APR.