About the Authors

Takaaki Daimon

Affiliations National Institute of Agrobiological Sciences, Tsukuba, Japan, Department of Agricultural and Environmental Biology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Toshinori Kozaki

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Ryusuke Niwa

Affiliation Initiative for the Promotion of Young Scientists' Independent Research, Graduate School of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Japan

Isao Kobayashi

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Kenjiro Furuta

Address Current address: Department of Life Science and Biotechnology, Faculty of Life and Environmental Science, Shimane University, Matsue, Japan

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Toshiki Namiki

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Keiro Uchino

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Yutaka Banno

Affiliation Institute of Genetic Resources, Faculty of Agriculture, Kyushu University Graduate School, Fukuoka, Japan

Susumu Katsuma

Affiliation Department of Agricultural and Environmental Biology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Toshiki Tamura

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Kazuei Mita

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Hideki Sezutsu

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Masayoshi Nakayama

Affiliation Institute of Floricultural Sciences, National Agriculture and Food Research Organization, Tsukuba, Japan

Kyo Itoyama

Affiliation School of Agriculture, Meiji University, Kawasaki, Japan

Toru Shimada

Affiliation Department of Agricultural and Environmental Biology, Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, Japan

Tetsuro Shinoda

shinoda@affrc.go.jp

Affiliation National Institute of Agrobiological Sciences, Tsukuba, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: T Daimon, T Kozaki, R Niwa, S Katsuma, H Sezutsu, T Shimada, T Shinoda. Performed the experiments: T Daimon, T Kozaki, R Niwa, I Kobayashi, K Furuta, T Namiki, K Uchino, M Nakayama, K Itoyama, T Shinoda. Analyzed the data: T Daimon, T Kozaki, R Niwa, T Namiki, T Shinoda. Contributed reagents/materials/analysis tools: I Kobayashi, K Uchino, Y Banno, T Tamura, K Mita, H Sezutsu. Wrote the paper: T Daimon, T Kozaki, R Niwa, K Furuta, T Shinoda.