About the Authors

Michael P. Heaton

mike.heaton@ars.usda.gov

Affiliation U.S. Meat Animal Research Center, Agriculture Research Service, United States Department of Agriculture, Clay Center, Nebraska, United States of America

Michael L. Clawson

Affiliation U.S. Meat Animal Research Center, Agriculture Research Service, United States Department of Agriculture, Clay Center, Nebraska, United States of America

Carol G. Chitko-Mckown

Affiliation U.S. Meat Animal Research Center, Agriculture Research Service, United States Department of Agriculture, Clay Center, Nebraska, United States of America

Kreg A. Leymaster

Affiliation U.S. Meat Animal Research Center, Agriculture Research Service, United States Department of Agriculture, Clay Center, Nebraska, United States of America

Timothy P. L. Smith

Affiliation U.S. Meat Animal Research Center, Agriculture Research Service, United States Department of Agriculture, Clay Center, Nebraska, United States of America

Gregory P. Harhay

Affiliation U.S. Meat Animal Research Center, Agriculture Research Service, United States Department of Agriculture, Clay Center, Nebraska, United States of America

Stephen N. White

Affiliation Animal Disease Research Unit, Agriculture Research Service, United States Department of Agriculture, Pullman, Washington, United States of America

Lynn M. Herrmann-Hoesing

Affiliation Animal Disease Research Unit, Agriculture Research Service, United States Department of Agriculture, Pullman, Washington, United States of America

Michelle R. Mousel

Affiliation U.S. Sheep Experiment Station, Agriculture Research Service, United States Department of Agriculture, Dubois, Idaho, United States of America

Gregory S. Lewis

Affiliation U.S. Sheep Experiment Station, Agriculture Research Service, United States Department of Agriculture, Dubois, Idaho, United States of America

Theodore S. Kalbfleisch

Affiliation Department of Biochemistry and Molecular Biology, School of Medicine, University of Louisville, Louisville, Kentucky, United States of America

James E. Keen

Affiliation Great Plains Veterinary Educational Center, University of Nebraska, Clay Center, Nebraska, United States of America

William W. Laegreid

Affiliation Department of Pathobiology, University of Illinois at Urbana-Champaign, Urbana, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MPH MLC CGC-M KAL TPLS SNW LMH-H MRM JEK WWL. Performed the experiments: MPH MLC CGC-M KAL TPLS SNW LMH-H MRM JEK WWL. Analyzed the data: MPH MLC CGC-M KAL TPLS SNW LMH-H MRM. Contributed reagents/materials/analysis tools: MPH MLC CGC-M KAL TPLS GPH GSL SNW LMH-H MRM TSK JEK WWL. Wrote the paper: MPH MLC CGC-M KAL TPLS GPH SNW LMH-H MRM GSL TSK JEK WWL.