About the Authors

Nicholas A. Johnson

Address Current address: Google, Mountainview, California, United States of America

Affiliation Department of Statistics, Stanford University, Stanford, California, United States of America

Marc A. Coram

Affiliation Department of Health Research and Policy, Stanford University School of Medicine, Stanford, California, United States of America

Mark D. Shriver

Affiliation Department of Anthropology, Pennsylvania State University, University Park, Pennsylvania, United States of America

Isabelle Romieu

Address Current address: International Agency for Research on Cancer, Lyon, France

Affiliation National Institute of Public Health, Cuernavaca, Mexico

Gregory S. Barsh

gbarsh@hudsonalpha.org (GB); huatang@stanford.edu (HT)

Affiliations Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America, HudsonAlpha Institute for Biotechnology, Huntsville, Alabama, United States of America

Stephanie J. London

Affiliations Epidemiology Branch, National Institute of Environmental Health Sciences, National Institutes of Health, Department of Health and Human Services, Research Triangle Park, North Carolina, United States of America, Laboratory of Respiratory Biology, National Institute of Environmental Health Sciences, National Institutes of Health, Department of Health and Human Services, Research Triangle Park, North Carolina, United States of America

Hua Tang

gbarsh@hudsonalpha.org (GB); huatang@stanford.edu (HT)

Affiliation Department of Genetics, Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: NAJ MAC GSB HT. Performed the experiments: NAJ MAC HT. Analyzed the data: NAJ MAC HT. Contributed reagents/materials/analysis tools: IR MDS SJL. Wrote the paper: GSB MAC SJL MDS HT.