About the Authors

Seongjin Seo

Address Current address: Department of Ophthalmology and Visual Sciences, University of Iowa College of Medicine, Iowa City, Iowa, United States of America

Affiliations Department of Pediatrics, University of Iowa College of Medicine, Iowa City, Iowa, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Qihong Zhang

Affiliations Department of Pediatrics, University of Iowa College of Medicine, Iowa City, Iowa, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Kevin Bugge

Affiliations Department of Pediatrics, University of Iowa College of Medicine, Iowa City, Iowa, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

David K. Breslow

Affiliation Department of Molecular and Cellular Physiology, Stanford University School of Medicine, Stanford, California, United States of America

Charles C. Searby

Affiliations Department of Pediatrics, University of Iowa College of Medicine, Iowa City, Iowa, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Maxence V. Nachury

Affiliation Department of Molecular and Cellular Physiology, Stanford University School of Medicine, Stanford, California, United States of America

Val C. Sheffield

val-sheffield@uiowa.edu

Affiliations Department of Pediatrics, University of Iowa College of Medicine, Iowa City, Iowa, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SS MVN VCS. Performed the experiments: SS QZ KB DKB CCS. Analyzed the data: SS MVN VCS. Wrote the paper: SS VCS.