About the Authors

Craig S. Nowell

Address Current address: EPFL SV ISREC UPRAD, SV 2533 (Batiment SV), Lausanne, Switzerland

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Nicholas Bredenkamp

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Stéphanie Tetélin

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Xin Jin

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Christin Tischner

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Harsh Vaidya

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Julie M. Sheridan

Address Current address: Walter and Eliza Hall Institute, Parkville, Australia

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Frances Hogg Stenhouse

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Raphaela Heussen

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Andrew J. H. Smith

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

C. Clare Blackburn

c.blackburn@ed.ac.uk

Affiliation MRC Centre for Regenerative Medicine, Institute for Stem Cell Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CSN NB JMS AJHS CCB. Performed the experiments: CSN NB ST XJ HV CT FHS RH. Analyzed the data: CSN NB ST XJ HV CT JMS FHS RH AJHS CCB. Contributed reagents/materials/analysis tools: CSN AJHS CCB. Wrote the paper: CSN CCB.