About the Authors

Heather A. Lawson

lawsonh@pcg.wustl.edu

Affiliation Washington University in St Louis, St Louis, Missouri, United States of America

Janet E. Cady

Affiliation Washington University in St Louis, St Louis, Missouri, United States of America

Charlyn Partridge

Affiliation Washington University in St Louis, St Louis, Missouri, United States of America

Jason B. Wolf

Affiliation University of Bath, Bath, United Kingdom

Clay F. Semenkovich

Affiliation Washington University in St Louis, St Louis, Missouri, United States of America

James M. Cheverud

Affiliation Washington University in St Louis, St Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HAL JMC. Performed the experiments: HAL JMC. Analyzed the data: HAL JEC JMC. Contributed reagents/materials/analysis tools: CFS. Wrote the paper: HAL. Prepared expression data for analysis: CP. Developed equations for imputing imprinting: JBW JMC. Assisted in editing manuscript: JBW CFS JMC.