About the Authors

Chris Cotsapas

Contributed equally to this work with: Chris Cotsapas, Benjamin F. Voight

Affiliations Center For Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Department of Medicine, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Neurology, Yale University School of Medicine, New Haven, Connecticut, United States of America, Department of Genetics, Yale University School of Medicine, New Haven, Connecticut, United States of America

Benjamin F. Voight

Contributed equally to this work with: Chris Cotsapas, Benjamin F. Voight

Affiliations Center For Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Department of Medicine, Harvard Medical School, Boston, Massachusetts, United States of America

Elizabeth Rossin

Affiliations Center For Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Department of Medicine, Harvard Medical School, Boston, Massachusetts, United States of America, Health Science and Technology MD Program, Harvard University and Massachusetts Institute of Technology, Boston, Massachusetts, United States of America, Harvard Biological and Biomedical Sciences Program, Harvard University, Boston, Massachusetts, United States of America

Kasper Lage

Affiliations Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Pediatric Surgical Research Laboratories, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Center for Biological Sequence Analysis, Department of Systems Biology, Technical University of Denmark, Lyngby, Denmark

Benjamin M. Neale

Affiliations Center For Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Department of Medicine, Harvard Medical School, Boston, Massachusetts, United States of America, Analytical and Translational Genetics Unity, Massachusetts General Hospital, Boston, Massachusetts, United States of America

Chris Wallace

Affiliation Juvenile Diabetes Research Foundation/Wellcome Trust Diabetes and Inflammation Laboratory, Department of Medical Genetics, Cambridge Institute for Medical Research, University of Cambridge, Cambridge, United Kingdom

Gonçalo R. Abecasis

Affiliation Center for Statistical Genetics, University of Michigan, Ann Arbor, Massachusetts, United States of America

Jeffrey C. Barrett

Affiliation Human Genetics, Wellcome Trust Sanger Institute, Cambridge, United Kingdom

Timothy Behrens

Affiliation Genentech, South San Francisco, California, United States of America

Judy Cho

Affiliations Department of Genetics, Yale University School of Medicine, New Haven, Connecticut, United States of America, Departments of Medicine and Genetics, Yale University School of Medicine, New Haven, Connecticut, United States of America

Philip L. De Jager

Affiliations Department of Medicine, Harvard Medical School, Boston, Massachusetts, United States of America, Department of Neurology, Brigham and Women's Hospital, Boston, Massachusetts, United States of America

James T. Elder

Affiliation Department of Dermatology, University of Michigan, Ann Arbor, Michigan, United States of America

Robert R. Graham

Affiliation Genentech, South San Francisco, California, United States of America

Peter Gregersen

Affiliation Feinstein Institute for Medical Research, North Shore-Long Island Jewish Health System, Manhasset, New York, United States of America

Lars Klareskog

Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institutet, Stockholm, Sweden

Katherine A. Siminovitch

Affiliation Samuel Lunenfeld Research Institute, Mount Sinai Hospital, Toronto, Canada

David A. van Heel

Affiliation Blizard Institute, The London School of Medicine and Dentistry, London, United Kingdom

Cisca Wijmenga

Affiliation Department of Genetics, University Medical Center Groningen and Groningen University, Groningen, The Netherlands

Jane Worthington

Affiliation Arthritis Research UK Epidemiology Unit, School of Translational Medicine, Manchester Academic Health Sciences Centre, University of Manchester, Manchester, United Kingdom

John A. Todd

Affiliation Juvenile Diabetes Research Foundation/Wellcome Trust Diabetes and Inflammation Laboratory, Department of Medical Genetics, Cambridge Institute for Medical Research, University of Cambridge, Cambridge, United Kingdom

David A. Hafler

Affiliation Department of Neurology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Stephen S. Rich

Affiliation Center for Public Health Genomics, University of Virginia, Charlottesville, Virginia, United States of America

Mark J. Daly

mjdaly@atgu.mgh.harvard.edu

Affiliations Center For Human Genetic Research, Massachusetts General Hospital, Boston, Massachusetts, United States of America, Medical and Population Genetics, Broad Institute of MIT and Harvard, Cambridge, Massachusetts, United States of America, Department of Medicine, Harvard Medical School, Boston, Massachusetts, United States of America, Analytical and Translational Genetics Unity, Massachusetts General Hospital, Boston, Massachusetts, United States of America

on behalf of the FOCiS Network of Consortia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: C Cotsapas, BF Voight, DA Hafler, SS Rich, MJ Daly. Performed the experiments: C Cotsapas, BF Voight, E Rossin, BM Neale, MJ Daly. Analyzed the data: C Cotsapas, BF Voight, E Rossin, K Lage, MJ Daly. Contributed reagents/materials/analysis tools: BF Voight, K Lage, BM Neale, C Wallace, GR Abecasis, JC Barrett, T Behrens, J Cho, PL De Jager, JT Elder, RR Graham, P Gregersen, L Klareskog, KA Siminovitch, DA van Heel, C Wijmenga, J Worthington, JA Todd, DA Hafler, SS Rich, MJ Daly. Wrote the paper: C Cotsapas, BF Voight, JA Todd, DA Hafler, SS Rich, MJ Daly.