About the Authors

Adam N. Famoso

Address Current address: Pioneer Hi-Bred International, Johnston, Iowa, United States of America

Affiliation Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America

Keyan Zhao

Address Current address: Department of Internal Medicine, University of Iowa, Iowa City, Iowa, United States of America

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Randy T. Clark

Affiliation Robert W. Holley Center for Agriculture and Health, Agricultural Research Service, US Department of Agriculture, Cornell University, Ithaca, New York, United States of America

Chih-Wei Tung

Affiliation Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America

Mark H. Wright

Affiliations Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America, Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Carlos Bustamante

Address Current address: Department of Genetics, Stanford University, Palo Alto, California, United States of America

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Leon V. Kochian

Affiliation Robert W. Holley Center for Agriculture and Health, Agricultural Research Service, US Department of Agriculture, Cornell University, Ithaca, New York, United States of America

Susan R. McCouch

srm4@cornell.edu

Affiliation Department of Plant Breeding and Genetics, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ANF RJC LVK SRM. Performed the experiments: ANF RTC. Analyzed the data: ANF KZ C-WT CB SRM. Wrote the paper: ANF KZ RTC C-WT MHW CB LVK SRM.