About the Authors

Joseph C. Maranville

Contributed equally to this work with: Joseph C. Maranville, Francesca Luca

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Francesca Luca

Contributed equally to this work with: Joseph C. Maranville, Francesca Luca

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Allison L. Richards

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Xiaoquan Wen

Affiliation Department of Statistics, The University of Chicago, Chicago, Illinois, United States of America

David B. Witonsky

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Shaneen Baxter

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Matthew Stephens

stephens@galton.uchicago.edu (MS); dirienzo@bsd.uchicago.edu (ADR)

Affiliations Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America, Department of Statistics, The University of Chicago, Chicago, Illinois, United States of America

Anna Di Rienzo

stephens@galton.uchicago.edu (MS); dirienzo@bsd.uchicago.edu (ADR)

Affiliation Department of Human Genetics, The University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JCM FL ADR. Performed the experiments: JCM FL ALR SB. Analyzed the data: JCM FL. Contributed reagents/materials/analysis tools: MS XW DBW. Wrote the paper: JCM FL ADR MS.