About the Authors

Barbara G. Mellone

barbara.mellone@uconn.edu (BGM); karpen@fruitfly.org (GHK)

Affiliation Department of Molecular and Cell Biology, University of Connecticut, Storrs, Connecticut, United States of America

Kathryn J. Grive

Contributed equally to this work with: Kathryn J. Grive, Vladimir Shteyn

Address Current address: Department of Molecular Biology, Cell Biology, and Biochemistry, Brown University, Providence, Rhode Island, United States of America

Affiliation Department of Molecular and Cell Biology, University of Connecticut, Storrs, Connecticut, United States of America

Vladimir Shteyn

Contributed equally to this work with: Kathryn J. Grive, Vladimir Shteyn

Address Current address: Department of Cell Biology, Yale University School of Medicine, New Haven, Connecticut, United States of America

Affiliations Department of Genome Dynamics, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America

Sarion R. Bowers

Affiliation Department of Molecular and Cell Biology, University of Connecticut, Storrs, Connecticut, United States of America

Isaac Oderberg

Address Current address: Department of Biology, Massachusetts Institute of Technology, Cambridge, Massachusetts, United States of America

Affiliations Department of Genome Dynamics, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America

Gary H. Karpen

barbara.mellone@uconn.edu (BGM); karpen@fruitfly.org (GHK)

Affiliations Department of Genome Dynamics, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BGM GHK. Performed the experiments: BGM KJG VS SRB. Analyzed the data: BGM KJG IO GHK. Contributed reagents/materials/analysis tools: BGM VS IO GHK. Wrote the paper: BGM GHK.