About the Authors

Chris Spencer, Damjan Vukcevic, Peter Donnelly
Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Eliana Hechter, Peter Donnelly
Department of Statistics, University of Oxford, Oxford, United Kingdom

Corresponding Authors

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CS DV PD. Performed the experiments: CS EH DV. Analyzed the data: CS EH DV PD. Contributed reagents/materials/analysis tools: CS. Wrote the paper: CS PD.