About the Authors

Chris Spencer

chris.spencer@well.ox.ac.uk (CS); peter.donnelly@well.ox.ac.uk (PD)

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Eliana Hechter

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

Damjan Vukcevic

Affiliation Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Peter Donnelly

chris.spencer@well.ox.ac.uk (CS); peter.donnelly@well.ox.ac.uk (PD)

Affiliations Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom, Department of Statistics, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CS DV PD. Performed the experiments: CS EH DV. Analyzed the data: CS EH DV PD. Contributed reagents/materials/analysis tools: CS. Wrote the paper: CS PD.