About the Authors

Sharon A. Chung

sharon.chung@ucsf.edu (SAC); lindsey.criswell@ucsf.edu (LAC)

Affiliation Rosalind Russell Medical Research Center for Arthritis, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Kimberly E. Taylor

Affiliation Rosalind Russell Medical Research Center for Arthritis, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Robert R. Graham

Affiliation ITGR Human Genetics, Genentech, South San Francisco, California, United States of America

Joanne Nititham

Affiliation Rosalind Russell Medical Research Center for Arthritis, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Annette T. Lee

Affiliation Robert S. Boas Center for Genomics and Human Genetics, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

Ward A. Ortmann

Affiliation ITGR Human Genetics, Genentech, South San Francisco, California, United States of America

Chaim O. Jacob

Affiliation Department of Medicine, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

Marta E. Alarcón-Riquelme

Affiliations Department of Genetics and Pathology, Uppsala University, Uppsala, Sweden, Andalusian Center for Genomics and Oncological Research, Pfizer-University of Granada-Junta de Andalucía, Granada, Spain, Arthritis and Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Betty P. Tsao

Affiliation Department of Medicine, University of California Los Angeles, Los Angeles, California, United States of America

John B. Harley

Affiliation US Department of Veterans Affairs Medical Center and Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

Patrick M. Gaffney

Affiliation Arthritis and Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Kathy L. Moser

Affiliation Arthritis and Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

SLEGEN

For membership of The International Consortium on the Genetics of Systemic Lupus Erythematosus please see the Acknowledgments.

Michelle Petri

Affiliation Division of Rheumatology, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

F. Yesim Demirci

Affiliation Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

M. Ilyas Kamboh

Affiliation Department of Human Genetics, Graduate School of Public Health, University of Pittsburgh, Pittsburgh, Pennsylvania, United States of America

Susan Manzi

Affiliation Allegheny-Singer Research Institute, Pittsburgh, Pennsylvania, United States of America

Peter K. Gregersen

Affiliation Robert S. Boas Center for Genomics and Human Genetics, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

Carl D. Langefeld

Affiliation Department of Biostatistical Sciences, Wake Forest University Health Sciences, Wake Forest, North Carolina, United States of America

Timothy W. Behrens

Affiliation ITGR Human Genetics, Genentech, South San Francisco, California, United States of America

Lindsey A. Criswell

sharon.chung@ucsf.edu (SAC); lindsey.criswell@ucsf.edu (LAC)

Affiliation Rosalind Russell Medical Research Center for Arthritis, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SAC LAC. Performed the experiments: SAC KET JN. Analyzed the data: SAC KET. Contributed reagents/materials/analysis tools: SAC KET RRG JN ATL WAO COJ MEAR BPT JBH PMG KLM MP FYD MIK SM PKG CDL TWB LAC. Wrote the paper: SAC LAC.