About the Authors

Benjamin Callahan

benjc@stanford.edu

Affiliation Department of Applied Physics, Stanford University, Stanford, California, United States of America

Richard A. Neher

Address Current address: Max-Planck-Institute for Developmental Biology, Tübingen, Germany

Affiliation Kavli Institute for Theoretical Physics, University of California Santa Barbara, Santa Barbara, California, United States of America

Doris Bachtrog

Affiliation Department of Integrative Biology and Center for Theoretical Evolutionary Genomics, University of California Berkeley, Berkeley, California, United States of America

Peter Andolfatto

Affiliation Department of Ecology and Evolutionary Biology and the Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Boris I. Shraiman

Affiliations Kavli Institute for Theoretical Physics, University of California Santa Barbara, Santa Barbara, California, United States of America, Department of Physics, University of California Santa Barbara, Santa Barbara, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BC BIS. Performed the experiments: DB PA. Analyzed the data: BC RAN. Contributed reagents/materials/analysis tools: BC DB PA. Wrote the paper: BC RAN PA BIS.