About the Authors

Kimberly E. Taylor

Affiliation Rosalind Russell Medical Research Center for Arthritis, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Sharon A. Chung

Affiliation Rosalind Russell Medical Research Center for Arthritis, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Robert R. Graham

Affiliation ITGR Human Genetics, Genentech, South San Francisco, California, United States of America

Ward A. Ortmann

Affiliation ITGR Human Genetics, Genentech, South San Francisco, California, United States of America

Annette T. Lee

Affiliation Robert S. Boas Center for Genomics and Human Genetics, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

Carl D. Langefeld

Affiliation Department of Biostatistical Sciences, Wake Forest University Health Sciences, Wake Forest, North Carolina, United States of America

Chaim O. Jacob

Affiliation Department of Medicine, Keck School of Medicine, University of Southern California, Los Angeles, California, United States of America

M. Ilyas Kamboh

Affiliation Department of Human Genetics, University of Pittsburgh Graduate School of Public Health, Pittsburgh, Pennsylvania, United States of America

Marta E. Alarcón-Riquelme

Affiliations Arthritis and Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America, Center for Genomics and Oncological Research, Pfizer-University of Granada-Junta de Andalucia, Granada, Spain

Betty P. Tsao

Affiliation Department of Medicine, University of California Los Angeles, Los Angeles, California, United States of America

Kathy L. Moser

Affiliation Arthritis and Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

Patrick M. Gaffney

Affiliation Arthritis and Immunology Research Program, Oklahoma Medical Research Foundation, Oklahoma City, Oklahoma, United States of America

John B. Harley

Affiliations U.S. Department of Veterans Affairs Medical Center Cincinnati, Ohio, United States of America, Cincinnati Children's Hospital Medical Center, Cincinnati, Ohio, United States of America

Michelle Petri

Affiliation Division of Rheumatology, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Susan Manzi

Affiliation Allegheny-Singer Research Institute, Pittsburgh, Pennsylvania, United States of America

Peter K. Gregersen

Affiliation Robert S. Boas Center for Genomics and Human Genetics, Feinstein Institute for Medical Research, Manhasset, New York, United States of America

Timothy W. Behrens

Affiliation ITGR Human Genetics, Genentech, South San Francisco, California, United States of America

Lindsey A. Criswell

lindsey.criswell@ucsf.edu

Affiliation Rosalind Russell Medical Research Center for Arthritis, Department of Medicine, University of California San Francisco, San Francisco, California, United States of America

Competing Interests

RRG and TWB are full-time employees of Genentech.

Author Contributions

Conceived and designed the experiments: KET SAC RRG PKG TWB LAC. Analyzed the data: KET CDL. Contributed reagents/materials/analysis tools: RRG WAO COJ PKG TWB. Wrote the paper: KET. Performed genotyping: ATL. Recruited subjects and collected data: MIK MEAR BPT KLM PMG JBH SM MP.