About the Authors

Kerry N. Veth

Affiliation Department of Cell Biology, Neurobiology, and Anatomy, Medical College of Wisconsin, Milwaukee, Wisconsin, United States of America

Jason R. Willer

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Ross F. Collery

Affiliation Department of Cell Biology, Neurobiology, and Anatomy, Medical College of Wisconsin, Milwaukee, Wisconsin, United States of America

Matthew P. Gray

Affiliation Department of Cell Biology, Neurobiology, and Anatomy, Medical College of Wisconsin, Milwaukee, Wisconsin, United States of America

Gregory B. Willer

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Daniel S. Wagner

Affiliation Department of Biochemistry and Cell Biology, Rice University, Houston, Texas, United States of America

Mary C. Mullins

Affiliation Department of Cell and Developmental Biology, University of Pennsylvania Medical School, Philadelphia, Pennsylvania, United States of America

Ava J. Udvadia

Affiliation Department of Biological Sciences, University of Wisconsin–Milwaukee, Milwaukee, Wisconsin, United States of America

Richard S. Smith

Affiliation Howard Hughes Medical Institute, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Simon W. M. John

Affiliation Howard Hughes Medical Institute, The Jackson Laboratory, Bar Harbor, Maine, United States of America

Ronald G. Gregg

Affiliation Department of Biochemistry and Molecular Biology, University of Louisville, Louisville, Kentucky, United States of America

Brian A. Link

blink@mcw.edu

Affiliation Department of Cell Biology, Neurobiology, and Anatomy, Medical College of Wisconsin, Milwaukee, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KNV RFC SWMJ RGG BAL. Performed the experiments: KNV JRW RFC MPG GBW RSS SWMJ BAL. Analyzed the data: KNV JRW RFC GBW RSS SWMJ RGG BAL. Contributed reagents/materials/analysis tools: KNV JRW RFC DSW MCM AJU RGG BAL. Wrote the paper: KNV BAL.