About the Authors

Justin K. Hines

Contributed equally to this work with: Justin K. Hines, Xiaomo Li

Affiliation Department of Biochemistry, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Xiaomo Li

Contributed equally to this work with: Justin K. Hines, Xiaomo Li

Affiliation Department of Molecular Pharmacology and Biological Chemistry, Feinberg School of Medicine, Northwestern University, Chicago, Illinois, United States of America

Zhiqiang Du

Affiliation Department of Molecular Pharmacology and Biological Chemistry, Feinberg School of Medicine, Northwestern University, Chicago, Illinois, United States of America

Takashi Higurashi

Address Current address: EVEC, Sapporo, Japan

Affiliation Department of Biochemistry, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Liming Li

ecraig@wisc.edu (EAC); limingli@northwestern.edu (LL)

Affiliation Department of Molecular Pharmacology and Biological Chemistry, Feinberg School of Medicine, Northwestern University, Chicago, Illinois, United States of America

Elizabeth A. Craig

ecraig@wisc.edu (EAC); limingli@northwestern.edu (LL)

Affiliation Department of Biochemistry, University of Wisconsin–Madison, Madison, Wisconsin, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JKH XL ZD TH LL EAC. Performed the experiments: JKH XL ZD TH. Analyzed the data: JKH XL ZD TH LL EAC. Wrote the paper: JKH XL LL EAC.