About the Authors

Margaret L. Hoang

Affiliations Howard Hughes Medical Institute and Department of Embryology, Carnegie Institution, Baltimore, Maryland, United States of America, Department of Biology, Johns Hopkins University, Baltimore, Maryland, United States of America

Frederick J. Tan

Address Current address: Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America

Affiliation Howard Hughes Medical Institute and Department of Embryology, Carnegie Institution, Baltimore, Maryland, United States of America

David C. Lai

Address Current address: University of Maryland, College Park, Maryland, United States of America

Affiliation Baltimore Polytechnic Institute, Ingenuity Program, Baltimore, Maryland, United States of America

Sue E. Celniker

Affiliation Life Sciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Roger A. Hoskins

Affiliation Life Sciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Maitreya J. Dunham

Affiliation Department of Genome Sciences, University of Washington, Seattle, Washington, United States of America

Yixian Zheng

Affiliation Howard Hughes Medical Institute and Department of Embryology, Carnegie Institution, Baltimore, Maryland, United States of America

Douglas Koshland

koshland@berkeley.edu

Address Current address: Department of Molecular and Cell Biology, University of California Berkeley, Berkeley, California, United States of America

Affiliation Howard Hughes Medical Institute and Department of Embryology, Carnegie Institution, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MLH FJT DK. Performed the experiments: MLH FJT DCL. Analyzed the data: MLH FJT DK. Contributed reagents/materials/analysis tools: MLH FJT SEC RAH MJD YZ. Wrote the paper: MLH DK.