About the Authors

Oscar M. Lancaster

Address Current address: Medical Research Council Laboratory for Molecular Cell Biology, Division of Life Sciences, University College London, London, United Kingdom

Affiliation The Wellcome Trust Centre for Cell Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Manuel Breuer

Affiliation The Wellcome Trust Centre for Cell Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

C. Fiona Cullen

Affiliation The Wellcome Trust Centre for Cell Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Takashi Ito

Affiliation Department of Biochemistry, Nagasaki University School of Medicine, Nagasaki, Japan

Hiroyuki Ohkura

h.ohkura@ed.ac.uk

Affiliation The Wellcome Trust Centre for Cell Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: OML MB HO. Performed the experiments: OML MB CFC. Analyzed the data: OML MB HO. Contributed reagents/materials/analysis tools: TI. Wrote the paper: OML HO.