About the Authors

Kouji Hirota

Affiliation CREST Research Project, Japan Science and Technology, Radiation Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Eiichiro Sonoda

Affiliation CREST Research Project, Japan Science and Technology, Radiation Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Takuo Kawamoto

Affiliation CREST Research Project, Japan Science and Technology, Radiation Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Akira Motegi

Affiliation CREST Research Project, Japan Science and Technology, Radiation Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Chikahide Masutani

Affiliation Solution-Oriented Research for Science and Technology (SORST), Japan Science and Technology Agency, Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan

Fumio Hanaoka

Address Current address: Department of Life Science, Faculty of Science, Gakushuin University, Tokyo, Japan

Affiliation Solution-Oriented Research for Science and Technology (SORST), Japan Science and Technology Agency, Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan

Dávid Szüts

Affiliation St. George's, University of London, London, United Kingdom

Shigenori Iwai

Affiliation Division of Chemistry, Graduate School of Engineering Science, Osaka University, Osaka, Japan

Julian E. Sale

Affiliation Medical Research Council Laboratory of Molecular Biology, Cambridge, United Kingdom

Alan Lehmann

Affiliation Genome Damage and Stability Centre, University of Sussex, Brighton, United Kingdom

Shunichi Takeda

stakeda@rg.med.kyoto-u.ac.jp

Affiliation CREST Research Project, Japan Science and Technology, Radiation Genetics, Graduate School of Medicine, Kyoto University, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KH ES TK AM ST. Performed the experiments: KH ES TK AM. Analyzed the data: KH ES TK AM ST. Contributed reagents/materials/analysis tools: CM FH DS SI JES AL. Wrote the paper: KH ES TK AM ST.