About the Authors

Kenneth D. Whitney


Affiliation Department of Ecology and Evolutionary Biology, Rice University, Houston, Texas, United States of America

Theodore Garland Jr

Affiliation Department of Biology, University of California Riverside, Riverside, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: KDW TG. Contributed reagents/materials/analysis tools: TG. Wrote the paper: KDW. Conceived and designed the study: KDW.