About the Authors

Kenneth D. Whitney
Department of Ecology and Evolutionary Biology, Rice University, Houston, Texas, United States of America

Theodore Garland Jr
Department of Biology, University of California Riverside, Riverside, California, United States of America

Corresponding Author

kwhitney@rice.edu

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: KDW TG. Contributed reagents/materials/analysis tools: TG. Wrote the paper: KDW. Conceived and designed the study: KDW.