About the Authors

Yi Lu

Contributed equally to this work with: Yi Lu, David P. Dimasi

Affiliation Genetics and Population Health, Queensland Institute of Medical Research, Brisbane, Australia

David P. Dimasi

Contributed equally to this work with: Yi Lu, David P. Dimasi

Affiliation Department of Ophthalmology, Flinders University, Adelaide, Australia

Pirro G. Hysi

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London School of Medicine, St Thomas' Hospital, London, United Kingdom

Alex W. Hewitt

Affiliation Centre for Eye Research Australia, University of Melbourne, Royal Victorian Eye and Ear Hospital, Melbourne, Australia

Kathryn P. Burdon

Affiliation Department of Ophthalmology, Flinders University, Adelaide, Australia

Tze'Yo Toh

Affiliations Centre for Eye Research Australia, University of Melbourne, Royal Victorian Eye and Ear Hospital, Melbourne, Australia, University of Tasmania, the Eye Hospital, Launceston, Australia

Jonathan B. Ruddle

Affiliation Centre for Eye Research Australia, University of Melbourne, Royal Victorian Eye and Ear Hospital, Melbourne, Australia

Yi Ju Li

Affiliations Center for Human Genetics, Duke University Medical Center, Durham, North Carolina, United States of America, Department of Biostatistics and Bioinformatics, Duke University Medical Center, Durham, North Carolina, United States of America

Paul Mitchell

Affiliation Centre for Vision Research, Westmead Millennium Institute, University of Sydney, Sydney, Australia

Paul R. Healey

Affiliation Centre for Vision Research, Westmead Millennium Institute, University of Sydney, Sydney, Australia

Grant W. Montgomery

Affiliation Genetics and Population Health, Queensland Institute of Medical Research, Brisbane, Australia

Narelle Hansell

Affiliation Genetics and Population Health, Queensland Institute of Medical Research, Brisbane, Australia

Timothy D. Spector

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London School of Medicine, St Thomas' Hospital, London, United Kingdom

Nicholas G. Martin

Affiliation Genetics and Population Health, Queensland Institute of Medical Research, Brisbane, Australia

Terri L. Young

Affiliation Center for Human Genetics, Duke University Medical Center, Durham, North Carolina, United States of America

Christopher J. Hammond

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London School of Medicine, St Thomas' Hospital, London, United Kingdom

Stuart Macgregor

These authors also contributed equally to this work.

Affiliation Genetics and Population Health, Queensland Institute of Medical Research, Brisbane, Australia

Jamie E. Craig

jamie.craig@flinders.edu.au

These authors also contributed equally to this work.

Affiliation Department of Ophthalmology, Flinders University, Adelaide, Australia

David A. Mackey

These authors also contributed equally to this work.

Affiliations Centre for Eye Research Australia, University of Melbourne, Royal Victorian Eye and Ear Hospital, Melbourne, Australia, Lions Eye Institute, University of Western Australia, Centre for Ophthalmology and Visual Science, Perth, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TLY CJH SM JEC DAM. Performed the experiments: DPD PH KPB TT GWM. Analyzed the data: YL DPD. Contributed reagents/materials/analysis tools: YL PH AWH KPB TT JBR YJL PM PRH GWM NH TDS NGM TLY CJH. Wrote the paper: YL DPD AWH SM JEC DAM.