About the Authors

Ching-Yu Cheng

cycheng@jhsph.edu (C-YC); wkao@jhsph.edu (WHLK)

Affiliations Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America, Department of Ophthalmology, Taipei Veterans General Hospital, Taipei, Taiwan, Department of Ophthalmology, National Yang Ming University School of Medicine, Taipei, Taiwan

David Reich

Affiliations Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of Harvard and Massachusetts Institute of Technology, Boston, Massachusetts, United States of America

Tien Y. Wong

Affiliations Singapore Eye Research Institute and Singapore National Eye Centre, Singapore, Centre for Eye Research Australia, University of Melbourne, Melbourne, Australia

Ronald Klein

Affiliation Department of Ophthalmology and Visual Sciences, University of Wisconsin School of Medicine and Public Health, Madison, Wisconsin, United States of America

Barbara E. K. Klein

Affiliation Department of Ophthalmology and Visual Sciences, University of Wisconsin School of Medicine and Public Health, Madison, Wisconsin, United States of America

Nick Patterson

Affiliation Program in Medical and Population Genetics, Broad Institute of Harvard and Massachusetts Institute of Technology, Boston, Massachusetts, United States of America

Arti Tandon

Affiliations Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America, Program in Medical and Population Genetics, Broad Institute of Harvard and Massachusetts Institute of Technology, Boston, Massachusetts, United States of America

Man Li

Affiliation Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Eric Boerwinkle

Affiliation Human Genetics Center, University of Texas Health Science Center at Houston, Houston, Texas, United States of America

A. Richey Sharrett

Affiliations Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America, Welch Center for Prevention, Epidemiology, and Clinical Research, Johns Hopkins University, Baltimore, Maryland, United States of America

W. H. Linda Kao

cycheng@jhsph.edu (C-YC); wkao@jhsph.edu (WHLK)

Affiliations Department of Epidemiology, Johns Hopkins Bloomberg School of Public Health, Baltimore, Maryland, United States of America, Welch Center for Prevention, Epidemiology, and Clinical Research, Johns Hopkins University, Baltimore, Maryland, United States of America, Department of Medicine, Johns Hopkins University School of Medicine, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CYC DR WHLK. Performed the experiments: DR WHLK. Analyzed the data: CYC NP AT ML. Contributed reagents/materials/analysis tools: DR TYW RK BEKK NP AT EB ARS WHLK. Wrote the paper: CYC WHLK.