About the Authors

Jennifer D. Lewis

Affiliation Department of Cell and Systems Biology, University of Toronto, Toronto, Ontario, Canada

Ronald Wu

Affiliation Department of Cell and Systems Biology, University of Toronto, Toronto, Ontario, Canada

David S. Guttman

Contributed equally to this work with: David S. Guttman, Darrell Desveaux

Affiliations Department of Cell and Systems Biology, University of Toronto, Toronto, Ontario, Canada, Centre for the Analysis of Genome Evolution and Function, University of Toronto, Toronto, Ontario, Canada

Darrell Desveaux

Contributed equally to this work with: David S. Guttman, Darrell Desveaux

darrell.desveaux@utoronto.ca

Affiliations Department of Cell and Systems Biology, University of Toronto, Toronto, Ontario, Canada, Centre for the Analysis of Genome Evolution and Function, University of Toronto, Toronto, Ontario, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JDL DSG DD. Performed the experiments: JDL RW. Analyzed the data: JDL RW DSG DD. Contributed reagents/materials/analysis tools: JDL DSG DD. Wrote the paper: JDL DSG DD.