About the Authors

Dan L. Nicolae

Affiliations Department of Medicine, University of Chicago, Chicago, Illinois, United States of America, Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America, Department of Statistics, University of Chicago, Chicago, Illinois, United States of America

Eric Gamazon

Affiliation Department of Medicine, University of Chicago, Chicago, Illinois, United States of America

Wei Zhang

Affiliation Department of Medicine, University of Chicago, Chicago, Illinois, United States of America

Shiwei Duan

Address Current address: Singapore Institute for Clinical Sciences, Agency for Science, Technology and Research, Singapore

Affiliation Department of Medicine, University of Chicago, Chicago, Illinois, United States of America

M. Eileen Dolan

Affiliations Department of Medicine, University of Chicago, Chicago, Illinois, United States of America, Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Nancy J. Cox

ncox@bsd.uchicago.edu

Affiliations Department of Medicine, University of Chicago, Chicago, Illinois, United States of America, Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DLN MED NJC. Performed the experiments: EG. Analyzed the data: DLN EG WZ SD NJC. Contributed reagents/materials/analysis tools: EG WZ SD MED. Wrote the paper: DLN EG MED NJC.