About the Authors

Daniel L. Halligan

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Fiona Oliver

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Adam Eyre-Walker

Affiliation Centre for the Study of Evolution and School of Life Sciences, University of Sussex, Brighton, United Kingdom

Bettina Harr

Affiliation Max-Planck-Institute for Evolutionary Biology, Plön, Germany

Peter D. Keightley

keightley.plosgenet2010@gmail.com

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PDK. Performed the experiments: FO. Analyzed the data: DLH BH PDK. Contributed reagents/materials/analysis tools: BH. Wrote the paper: DLH AEW BH PDK.