About the Authors

Daniel L. Halligan, Fiona Oliver, Peter D. Keightley
Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Adam Eyre-Walker
Centre for the Study of Evolution and School of Life Sciences, University of Sussex, Brighton, United Kingdom

Bettina Harr
Max-Planck-Institute for Evolutionary Biology, Plön, Germany

Corresponding Author

keightley.plosgenet2010@gmail.com

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: PDK. Performed the experiments: FO. Analyzed the data: DLH BH PDK. Contributed reagents/materials/analysis tools: BH. Wrote the paper: DLH AEW BH PDK.