About the Authors

Liqin Cao

Contributed equally to this work with: Liqin Cao, Hiroshi Shitara

Affiliations Laboratory of Mouse Models for Human Heritable Diseases, The Tokyo Metropolitan Institute of Medical Science (Rinshoken), Tokyo, Japan, Technology and Development Team for Mammalian Cellular Dynamics, BioResource Center (BRC) RIKEN Tsukuba Institute, Ibaraki, Japan

Hiroshi Shitara

Contributed equally to this work with: Liqin Cao, Hiroshi Shitara

shitara-hr@igakuken.or.jp

Affiliation Laboratory of Mouse Models for Human Heritable Diseases, The Tokyo Metropolitan Institute of Medical Science (Rinshoken), Tokyo, Japan

Michihiko Sugimoto

Affiliation Technology and Development Team for Mammalian Cellular Dynamics, BioResource Center (BRC) RIKEN Tsukuba Institute, Ibaraki, Japan

Jun-Ichi Hayashi

Affiliation Graduate School of Life and Environmental Sciences, University of Tsukuba, Ibaraki, Japan

Kuniya Abe

Affiliation Technology and Development Team for Mammalian Cellular Dynamics, BioResource Center (BRC) RIKEN Tsukuba Institute, Ibaraki, Japan

Hiromichi Yonekawa

Affiliation Laboratory of Mouse Models for Human Heritable Diseases, The Tokyo Metropolitan Institute of Medical Science (Rinshoken), Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: LC HS JIH HY. Performed the experiments: LC HS MS. Analyzed the data: LC HS MS. Contributed reagents/materials/analysis tools: MS JIH KA HY. Wrote the paper: LC HS JIH KA HY.