About the Authors

Sara E. Beese

Affiliation Department of Biochemistry and Molecular Biology, Bloomberg School of Public Health, The Johns Hopkins University, Baltimore, Maryland, United States of America

Takahiro Negishi

Current address: Department of Integrated Biosciences, Graduate School of Frontier Sciences, University of Tokyo, Chiba, Japan

Affiliation Department of Biochemistry and Molecular Biology, Bloomberg School of Public Health, The Johns Hopkins University, Baltimore, Maryland, United States of America

David E. Levin

delevin@bu.edu

Affiliations Department of Biochemistry and Molecular Biology, Bloomberg School of Public Health, The Johns Hopkins University, Baltimore, Maryland, United States of America, Department of Molecular and Cell Biology, Boston University Goldman School of Dental Medicine, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SEB DEL. Performed the experiments: SEB TN DEL. Analyzed the data: SEB DEL. Contributed reagents/materials/analysis tools: SEB TN DEL. Wrote the paper: SEB DEL.