About the Authors

Sanzhen Liu

Affiliations Interdepartmental Genetics Graduate Program, Iowa State University, Ames, Iowa, United States of America, Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Cheng-Ting Yeh

Affiliations Center for Plant Genomics, Iowa State University, Ames, Iowa, United States of America, Department of Agronomy, Iowa State University, Ames, Iowa, United States of America

Tieming Ji

Affiliations Bioinformatics and Computational Biology Program, Iowa State University, Ames, Iowa, United States of America, Department of Statistics, Iowa State University, Ames, Iowa, United States of America

Kai Ying

Affiliations Interdepartmental Genetics Graduate Program, Iowa State University, Ames, Iowa, United States of America, Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America

Haiyan Wu

Address Current address: Roche Global Pharma Development Center, Shanghai, China

Affiliation Bioinformatics and Computational Biology Program, Iowa State University, Ames, Iowa, United States of America

Ho Man Tang

Affiliation Center for Plant Genomics, Iowa State University, Ames, Iowa, United States of America

Yan Fu

Affiliations Department of Agronomy, Iowa State University, Ames, Iowa, United States of America, Center for Carbon Capturing Crops, Iowa State University, Ames, Iowa, United States of America

Dan Nettleton

Affiliation Department of Statistics, Iowa State University, Ames, Iowa, United States of America

Patrick S. Schnable

schnable@iastate.edu

Affiliations Interdepartmental Genetics Graduate Program, Iowa State University, Ames, Iowa, United States of America, Department of Genetics, Development, and Cell Biology, Iowa State University, Ames, Iowa, United States of America, Center for Plant Genomics, Iowa State University, Ames, Iowa, United States of America, Department of Agronomy, Iowa State University, Ames, Iowa, United States of America, Bioinformatics and Computational Biology Program, Iowa State University, Ames, Iowa, United States of America, Center for Carbon Capturing Crops, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SL PSS. Performed the experiments: SL HMT. Analyzed the data: SL CTY TJ KY HW YF DN PSS. Contributed reagents/materials/analysis tools: SL CTY TJ KY HW DN PSS. Wrote the paper: SL CTY YF DN PSS.