About the Authors

Margaret C. W. Ho

Contributed equally to this work with: Margaret C. W. Ho, Holly Johnsen, Sara E. Goetz

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

Holly Johnsen

Contributed equally to this work with: Margaret C. W. Ho, Holly Johnsen, Sara E. Goetz

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

Sara E. Goetz

Contributed equally to this work with: Margaret C. W. Ho, Holly Johnsen, Sara E. Goetz

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

Benjamin J. Schiller

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

Esther Bae

Affiliation College of Osteopathic Medicine of the Pacific, Western University of Health Sciences, Pomona, California, United States of America

Diana A. Tran

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

Andrey S. Shur

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

John M. Allen

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

Christoph Rau

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

Welcome Bender

Affiliation Department of Biological Chemistry and Molecular Pharmacology, Harvard Medical School, Boston, Massachusetts, United States of America

William W. Fisher

Affiliation Berkeley Drosophila Genome Project, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Susan E. Celniker

Affiliation Berkeley Drosophila Genome Project, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Robert A. Drewell

drewell@hmc.edu

Affiliation Biology Department, Harvey Mudd College, Claremont, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MCWH HJ SEG BJS EB ASS JMA CR WB SEC RAD. Performed the experiments: MCWH HJ SEG BJS EB DAT ASS JMA CR WB WWF RAD. Analyzed the data: MCWH HJ SEG BJS EB DAT ASS JMA CR WB SEC RAD. Contributed reagents/materials/analysis tools: WWF SEC RAD. Wrote the paper: MCWH HJ SEG EB ASS RAD.