About the Authors

Tony Kwan

Contributed equally to this work with: Tony Kwan, Elin Grundberg

tony.kwan@mcgill.ca

Affiliations Department of Human Genetics, McGill University, Montréal, Canada, McGill University and Genome Québec Innovation Centre, Montréal, Canada

Elin Grundberg

Contributed equally to this work with: Tony Kwan, Elin Grundberg

Affiliations Department of Human Genetics, McGill University, Montréal, Canada, McGill University and Genome Québec Innovation Centre, Montréal, Canada

Vonda Koka

Affiliation McGill University and Genome Québec Innovation Centre, Montréal, Canada

Bing Ge

Affiliation McGill University and Genome Québec Innovation Centre, Montréal, Canada

Kevin C. L. Lam

Affiliation McGill University and Genome Québec Innovation Centre, Montréal, Canada

Christel Dias

Affiliation McGill University and Genome Québec Innovation Centre, Montréal, Canada

Andreas Kindmark

Affiliation Department of Medical Sciences, Uppsala University, Uppsala, Sweden

Hans Mallmin

Affiliation Department of Surgical Sciences, Uppsala University, Uppsala, Sweden

Östen Ljunggren

Affiliation Department of Medical Sciences, Uppsala University, Uppsala, Sweden

Fernando Rivadeneira

Affiliations Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands, Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands

Karol Estrada

Affiliation Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands

Joyce B. van Meurs

Affiliation Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands

Andre Uitterlinden

Affiliations Department of Internal Medicine, Erasmus MC, Rotterdam, The Netherlands, Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands

Magnus Karlsson

Affiliation Clinical and Molecular Osteoporosis Research Unit, Department of Clinical Science, Lund University and Department of Orthopaedics, Malmö University Hospital, Malmö, Sweden

Claes Ohlsson

Affiliation Center for Bone Research at the Sahlgrenska Academy, Department of Internal Medicine, Göteborg University, Gothenburg, Sweden

Dan Mellström

Affiliation Center for Bone Research at the Sahlgrenska Academy, Department of Internal Medicine, Göteborg University, Gothenburg, Sweden

Olle Nilsson

Affiliation Department of Surgical Sciences, Uppsala University, Uppsala, Sweden

Tomi Pastinen

Affiliations Department of Human Genetics, McGill University, Montréal, Canada, McGill University and Genome Québec Innovation Centre, Montréal, Canada

Jacek Majewski

Affiliations Department of Human Genetics, McGill University, Montréal, Canada, McGill University and Genome Québec Innovation Centre, Montréal, Canada

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK EG CD TP JM. Performed the experiments: TK EG VK BG KCLL CD AK HM. Analyzed the data: TK EG VK BG CD AK FR KE JBvM. Contributed reagents/materials/analysis tools: TK EG VK BG KCLL CD AK HM ÖL FR KE JBvM AU MK CO DM ON TP JM. Wrote the paper: TK EG.