About the Authors

Keith A. Hultman

Affiliation Department of Genetics, Washington University School of Medicine, St. Louis, Missouri, United States of America

Erine H. Budi

Affiliation Department of Biology, Institute for Stem Cell and Regenerative Medicine, University of Washington, Seattle, Washington, United States of America

Daniel C. Teasley

Affiliation Department of Genetics, Washington University School of Medicine, St. Louis, Missouri, United States of America

Andrew Y. Gottlieb

Affiliation Department of Genetics, Washington University School of Medicine, St. Louis, Missouri, United States of America

David M. Parichy

Affiliation Department of Biology, Institute for Stem Cell and Regenerative Medicine, University of Washington, Seattle, Washington, United States of America

Stephen L. Johnson

sjohnson@genetics.wustl.edu

Affiliation Department of Genetics, Washington University School of Medicine, St. Louis, Missouri, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KAH SLJ. Performed the experiments: KAH EHB DCT AYG. Analyzed the data: KAH EHB DMP SLJ. Contributed reagents/materials/analysis tools: KAH EHB DMP SLJ. Wrote the paper: KAH SLJ.