About the Authors

Alkes L. Price

Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Arti Tandon

Affiliations Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America, Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Nick Patterson

Affiliation Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Kathleen C. Barnes

Affiliation Johns Hopkins Allergy and Asthma Center, Division of Clinical Immunology, Department of Medicine, School of Medicine, Baltimore, Maryland, United States of America

Nicholas Rafaels

Affiliation Johns Hopkins Allergy and Asthma Center, Division of Clinical Immunology, Department of Medicine, School of Medicine, Baltimore, Maryland, United States of America

Ingo Ruczinski

Affiliation Department of Biostatistics, Johns Hopkins School of Public Health, Baltimore, Maryland, United States of America

Terri H. Beaty

Affiliation Department of Biostatistics, Johns Hopkins School of Public Health, Baltimore, Maryland, United States of America

Rasika Mathias

Affiliation Inherited Disease Research Branch, National Human Genome Research Institute, National Institutes of Health, Baltimore, Maryland, United States of America

David Reich

reich@genetics.med.harvard.edu (DR); myers@stats.ox.ac.uk (SM)

Affiliations Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America, Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Simon Myers

reich@genetics.med.harvard.edu (DR); myers@stats.ox.ac.uk (SM)

Affiliations Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America, Department of Statistics, Oxford University, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ALP NP DR SM. Performed the experiments: ALP AT NP KCB NR IR THB RM DR SM. Analyzed the data: ALP AT NP DR SM. Contributed reagents/materials/analysis tools: ALP NP KCB NR IR THB RM DR SM. Wrote the paper: ALP AT NP KCB NR IR THB RM DR SM.